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Palynological evidence for late Westphalian–early Stephanian vegetation change in the Dobrudzha Coalfield, NE Bulgaria

TATIANA KH. DIMITROVA^* and CHRISTOPHER J. CLEAL^,

^* Geological Institute, Bulgarian Academy of Sciences, G. Bonchev Street Block 24, 1113 Sofia, Bulgaria
Department of Biodiversity & Systematic Biology, National Museum Wales, Cathays Park, Cardiff CF10 3NP, UK

Author for correspondence: chris.cleal{at}museumwales.ac.uk

The Dobrudzha Coalfield in northeast Bulgaria has coal-bearing deposits ranging from latest Namurian to early Stephanian age (late Bashkirian to Moscovian age). Palynology of the coals in the upper Makedonka, Krupen and Gurkovo formations has been used to identify major changes in the vegetation during late Westphalian and early Stephanian times. The palynomorphs were grouped in two different ways (according to general morphotype and according to parent plant group) and stratigraphical trends in the two sets of groups used to identify vegetation change through the succession. Detrended Correspondence Analysis was also used to identify ecological relationships between the palynofloras. In the upper Makedonka and Krupen formations, lycophyte spores mostly comprise 15–25 % of the palynofloras in the thicker seams, but in the thinner coals they can represent up to 55 %. Of the other plant groups, ferns are the most abundant, representing 31–69 % in the thicker seams, 12–41 % in the thinner seams. This suggests that the arborescent lycophytes were mainly restricted to pioneer vegetation in these upland areas, and were replaced by ferns as the peat substrates became better developed, suggesting better-drained conditions. The thinner seams also have a noticeable component of sphenophyll spores, indicating significant areas of open conditions allowing colonization by these scrambling plants. In the Gurkovo Formation coals, lycophytes form an even smaller part of the palynological spectra, usually less than 10 % and in only one sample just over 25 %; fern spores make up 43–57 % of the palynofloras. This is in contrast to the palynofloras reported from contemporaneous clastic deposits in South Wales, most of which consist mostly of 34–60 % lycophyte spores and 14–34 % fern spores. Even more marked is the difference from the contemporaneous coals formed in lowland settings in the USA, which have mostly > 50 % lycophyte spores. This evidence suggests that the timing of the decline in abundance of arborescent lycophytes varied according to elevation above sea-level. In lowland coastal settings, the lycophytes remained dominant until middle–late Cantabrian times, but in more inland areas they were progressively replaced mainly by arborescent ferns during late Westphalian times.

Key Words: palynology • Carboniferous • Dobrudzha Coalfield • Bulgaria • vegetation change